Please email me (t.ezard [at] for access to pdfs or source code, although I’m trying to move as much of the latter as possible to figshare or similar.

For metrics and impact, see Google Scholar or Impact Story.

Names in bold types indicate lab members for the project.

  1. -Brombacher, A., Elder,  L.E., Hull, P.M., Wilson, P.A. & Ezard, T.H.G. (in press) Calibration of test diameter and area as proxies for body size in the planktonic foraminifer Globoconella puncticulata. J. Foram. Res.

  2. -Nicol-Harper A., Dooley C.A., Packman D., Mueller, M., Bijak, J., Hodgson, D.J., Townley S. & Ezard T.H.G. (2018) Inferring transient dynamics of human populations from non-normality metrics. Popul. Ecol., 1-12. doi: 10.1007/s10144-018-0620-y.


  1. -Edgar, K.M., Hull, P.M. & Ezard, T.H.G. (2017) Evolutionary history biases inferences of ecology and environment from δ13C but not δ18O values. Nature Comms doi: 10.1038/s41467-017-01154-7.

  2. -    R code and data compilations on FigShare.

  3. -Pigeon, G., Ezard, T.H.G., Festa-Bianchet, M., Coltman, D.W., & Pelletier, F. (2017) Fluctuating effects of genetic and plastic changes in body mass on population dynamics in a large herbivore. Ecology 98, 2456-2467. doi: 10.1002/ecy.1940

  4. -Brombacher, A., Wilson, P.A., Bailey, I. & Ezard, T.H.G. (2017) The breakdown of static and evolutionary allometries during climate upheaval. American Naturalist 190, 350-362. doi: 10.1086/692570

  5. -    Data available on Dryad.

  6. -Brombacher, A., Wilson, P.A. & Ezard, T.H.G. (2017) Calibration of the repeatability of foraminiferal test size and shape measures with recommendations for future use. Mar. Micropal. 133, 21-27. doi: 10.1016/j.marmicro.2017.05.003

  7. -McDonald, J.L., Franco, M., Townley, S., Ezard, T.H.G., Jelbert, K. & Hodgson, D.J. (2017) Divergent demographic strategies of plants in variable environments. Nature Ecol. & Evol. 1, 0029. doi:10.1038/s41559-016-0029.

  8. -Rillo, M.C., Whittaker, J., Ezard, T.H.G., Purvis, A., Henderson, A. S., Stukins, S. & Miller, C.G. (online). The unknown planktonic foraminiferal pioneer Henry A. Buckley and his collection at The Natural History Museum, London. J. Foram. Res.. doi: 10.1144/jmpaleo2016-020.


  1. -Cooke, R.S.C., Woodfine, T., Petretto, M. & Ezard, T.H.G. (2016) Resource partitioning between ungulate populations in arid environments. Ecol. Evol. 6, 6354-6365. doi: 10.1002/ece3.2218.

  2. -Ezard, T.H.G. & Purvis, A. (2016) Environmental changes define ecological limits to species richness and the mode of macroevolutionary competition. Ecology Letters. 19, 899-906. doi: 10.1111/ele.12626.

  3. -Ezard, T.H.G., Quental, T.B., & Benton, M.B. (2016) The challenges to inferring the regulators of biodiversity in deep time. Phil. Trans. Roy. Soc. B. 371, 20150216. doi:10.1098/rstb.2015.0216

  4. -    Introduction to our Theme Issue on The Regulators of Biodiversity in Deep Time.


  1. -Spake, R., Ezard, T.H.G., Martin, P.A., Newton, A.C. & Doncaster, C.P. (2015) A meta-analysis of functional group responses to forest recovery outside of the tropics. Cons. Biol. 29, 1695-1703. doi: 10.1111/cobi.12548.

  2. -Ezard, T.H.G., Edgar, K.M. & Hull, P.M. (2015) Environmental and biological controls on size-specific δ13C and δ18O in recent planktonic foraminifera. Paleooceanography 30, 151–173. doi: 10.1002/2014PA002735.

  3. -    R code and data compilations on FigShare.

  4. -Jørgensen, P.S., Barraquand, F., Bonhomme, V., Curran, T.J., Cieraad, E., Ezard T.H.G., Gheradi, L.A., Hayes, R.A., Poisot, T., Salguero-Gómez, R., DeSoto, L., Swartz, B., Talbot, J.M., Wee, B. & Zimmerman, N. (2015). Connecting people and ideas from around the world: Global innovation platforms for next-generation ecology and beyond. Ecosphere 6:art68. doi: 10.1890/ES14-00198.1.


  1. -Prizak, R., Ezard, T.H.G. & Hoyle, R.B. (2014). Fitness consequences of maternal and grandmaternal effects. Ecol. Evol. 4, 3139–3145. doi: 10.1002/ece3.1150.

  2. -     MATLAB simulation routines on figshare.

  3. -Barraquand, F. Ezard, T.H.G., Joergensen, P., Zimmerman, N., Chamberlain, S.A., Salguero-Gómez, R., Curran, T.J. & Poisot, T. (2014) Lack of quantitative training among early-career ecologists: a survey of the problem and potential solutions. PeerJ 2:e285. doi: 10.7717/peerj.285

  4. -    Preprint & data.

  5. -Ezard, T.H.G., Prizak, R. & Hoyle, R.B. (2014). The fitness implications of adaptation via phenotypic plasticity and maternal effects. Func. Ecol. 28, 693–701. doi: 10.1111/1365-2435.12207.

  6. -     MATLAB simulation routines on figshare.

  7. -Seddon, A.W.R., Mackay, A.W., Baker, A.G., 32 others, then Ezard, T.H.G. & then another 33 others (2014) Looking forward through the past. Identification of fifty priority research questions in palaeoecology. J. Ecol.. 102, 256-267. doi: 10.1111/1365-2745.12195.

  8. -Pearson, P.N. & Ezard, T.H.G. (2014) Evolution and speciation in the Eocene planktonic foraminifer Turborotalia. Paleobiology. 40, 130-143. doi: 10.1666/13004.

  9. -     Supporting material (data, movies, additional figures) on figshare.


  1. -Ezard, T.H.G., Thomas, G.H. & Purvis, A. (2013) Inclusion of a near-complete fossil record reveals speciation-related molecular evolution. Methods Ecol. Evol. 4, 745-753. doi: 10.1111/2041-210X.12089

  2. -      Supporting Information (all phylogenies & data) via figshare.

  3. -      Cover image.

         -      Daily Mail and Guardian Higher Education Network press coverage.

  1. -Aze, T., Ezard, T.H.G., Purvis, A., Coxall, H.K., Stewart, D.R.M., Wade, B.S. & Pearson, P.N. (2013). Identifying anagenesis and cladogenesis in the fossil record. Proc. Natl. Acad. Sci USA 110, E2946. doi: 10.1073/pnas.1208302110.

  2. -   Reply to Strotz & Allen

  3. -Townley, S. & Ezard, T.H.G. (2013). A G matrix analogue to capture the cumulative effects of non-genetic inheritance. J. Evol. Biol. 26, 1234-1243. doi: 10.1111/jeb.12089

  4. -    R script

  5. -Hoyle, R.B. & Ezard, T.H.G. (2012). The benefits of maternal effects in novel and in stable environments. J. Roy. Soc. Interface 9, 2403-2413. doi:10.1098/rsif.2012.0183

  6. -    Podcast for the Journal of Ecology.

  7. -    Green OA version

  8. -    MATLAB simulation routines on figshare.

  9. -Ezard, T.H.G., Pearson, P.N., Aze, T., & Purvis, A. (2012). The Meaning of Birth and Death (in Macroevolutionary Birth-Death Models). Biol. Lett. 8, 139-142. doi: 10.1098/rsbl.2011.0699

  10. -    On cover for “Models in Palaeontology” special issue.


  1. -Ezard, T.H.G., Aze, T., Pearson, P.N. & Purvis, A. (2011). Interplay between Changing Climate and Species’ Ecology Drives Macroevolutionary Dynamics. Science 332, 349-351. doi: 10.1126/science.1203060

  2. -    Faculty of 1000 “must-read”.

  3. -    Pecha Kucha talk.

  4. -Aze, T., Ezard, T.H.G., Purvis, A., Coxall, H.K., Stewart, D.R.M., Wade, B.S. & Pearson, P.N. (2011). A phylogeny of Cenozoic macroperforate planktonic foraminifera from fossil data. Biol. Rev. 86, 900-927. doi: 10.1111/j.1469-185X.2011.00178.x

  5. -    Charles Downie Award 2011 for “best & most significant” micropalaeontogical paper.

  6. -Ezard, T.H.G., Pearson, P.N. & Purvis, A. (2010). Algorithmic approaches to aid species delimitation in multidimensional morphospace. BMC Evol. Biol., 10: 175, doi: 10.1186/1471-2148- 10-175

  7. -Ezard, T.H.G., Bullock, J.M., Dalgleish, H.J., Millon, A., Pelletier, F., Ozgul, A. & Koons, D.N. (2010). Matrix models for a changeable world: the importance of transient dynamics in population management. J. Appl. Ecol. 47, 515-523. doi: 10.1111/j.1365-2664.2010.01801.x

  8. -    Supplement including MATLAB and R code

  9. -Ezard, T.H.G. & Coulson, T. (2010). How sensitive are elasticities of long-run stochastic growth to how environmental variability is modelled? Ecol. Model. 221, 191-200, doi: 10.1016/j.ecolmodel.2009.09.017


  1. -Ezard, T.H.G. & Purvis, A. (2009). paleoPhylo: free software to draw paleobiological phylogenies. Paleobiology 35, 460-464. doi: 10.1666/0094-8373-35.3.460

  2. -Ezard, T.H.G., Côté, S.D. & Pelletier, F. (2009). Eco-evolutionary dynamics: disentangling phenotypic, environmental and population fluctuations. Phil. Trans. R. Soc. Lond. B 364, 1491- 1498. doi: 10.1098/rstb.2009.0006

  3. -    Figures reproduced in a Science review on eco-evolutionary dynamics.

  4. -Ezard, T.H.G., Gaillard, J.-M., Crawley, M.J. & Coulson, T. (2008). Habitat dependence and correlations between elasticities of long-term growth rates. Am. Nat. 172: 424-430. doi: 10.1086/589897

  5. -    Bright Club Guildford reworking as a stand-up comedy set.

  6. -Coulson, T., Ezard, T.H.G., Pelletier, F., Stenseth, N.C., Pilkington, J. Pemberton, J.M., Kruuk, L.E.B., Clutton-Brock, T.H. & Crawley, M. (2008) Estimating the functional form for the density dependence from life history data. Ecology 89: 1664-1674. doi: 10.1890/07-1099.1

  7. -Becker, P.H., Ezard, T.H.G., Ludwigs, J.-D., Sauer-Gürth, H. & Wink, M. (2008) Population sex ratio shift from fledging to recruitment: consequences for demography in a philopatric seabird. Oikos 117: 60-68. doi: 10.1111/j.2007.0030-1299.16287.x


  1. -Ezard, T.H.G., Becker, P.H. & Coulson, T. (2007) Correlations between Age, Phenotypic Traits and Individual Contributions to Population Growth in Common Terns. Ecology 88: 2496-2504. doi: 10.1890/06-2020.1

  2. -Dittmann, T., Ezard, T.H.G. & Becker, P.H. (2007) Prospectors colony attendance is sex-specific and increases future recruitment chances in a seabird. Behav. Proc. 76: 198-205. doi: 10.1016/j.beproc.2007.05.002

  3. -Ezard, T.H.G., Becker, P.H. & Coulson, T. (2006). The Contributions of Age and Sex to Variation in Common Tern Population Growth Rate. J. Anim. Ecol. 75: 1379-1386. doi: 10.1111/j.1365- 2656.2006.01162.x

  4. -Ezard, T.H.G. & Travis, J.M.J. (2006). The Impact of Habitat Loss and Fragmentation on Genetic Drift and Fixation Time. Oikos 114: 376-376. doi: 10.1111/j.2006.0030-1299.14778.x

  5. -Travis, J.M.J. & Ezard, T.H.G., (2006). Habitat geometry, population viscosity and the rate of genetic drift. Ecol. Inf. 1, 153-161. doi: j.ecoinf.2006.03.003.